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Kobel, H. R., Loumont C., & Tinsley R. C. (1996).  The extant species. The Biology of Xenopus. Symposia of the Zoological Society of London 68.. 9-33 in Tinsley, R.C., and H.R. Kobel, eds. . Oxford, UK.: Clarendon Press.
Bogart, J. P. (1980).  Evolutionary implication of polyploidy in amphibians and reptiles. (LewisW H., Ed.).Polyploidy: biological relevance. XII, 341-378. New York & London: Plenum Press.
Blair,(ed.). W. F. (1972).  Evolution in the genus Bufo. Austin: University of Texas Press.
van der Meijden, A. (2004).  Evidence for a South African endemic clade of Ranid frogs. V Congresso Nazionale della Societas Herpetologica Italica.
Pethiyagoda, R., & Manamendra-Arachchi K. (1999).  Evaluating Sri-Lanka's amphibian diversity. Occasional Papers of the Wildlife Heritage Trust. 2,
Laurent, R. F. (1954).  Etude de quelques epèces méconnues du genre Ptychadena. . Annales du Musée Royal du Congo Belge. Sciences Zoologiques. Tervuren. 34, 7-34.
Thibaudeau, G., & Altig R. (1999).  Endotrophic Anurans: Development and Evolution. McDiarmid, Roy W. [Author], Altig, Ronald [Author], 1999.
Matschie, P. (1983).  Einege anscheinend neue Reptilien und Amphibien aus West-Afrika.. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin . 170-175.