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Tandy, R. (1972).  The Evolution of African Bufo.
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Stokes, S., Thomas D. S. G., & Washington R. (1997).  Multiple episodes of aridity in southern Africa since the last interglacial period. Nature. 388, 154-158.
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Goebel, A. M., Donnelly J. M., & Atz M. E. (1999).  PCR primers and amplification methods for 12S ribosomal DNA, the control region, cytochrome oxidase I, and cytochrome b in Bufonids and other frogs, and an overview of PCR primers which have amplified DNA in amphibians successfully. Molecular Phylogenetics and Evolution. 11, 163-199.
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