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Nuclear gene phylogeny of narrow-mouthed toads (Family: Microhylidae) and a discussion of competing hypotheses concerning their biogeographical origins. Molecular Phylogenetics and Evolution. 44, 1017-1030.(2007).
A previously unrecognized radiation of ranid frogs in Southern Africa revealed by nuclear and mitochondrial DNA sequences. Molecular Phylogenetics & Evolution. 37, 674-685.(2005).
Novel phylogenetic relationships of the enigmatic brevicipitine and scaphiophrynine toads as revealed by sequences from the nuclear Rag-1 gene. Proceedings of the Royal Society of London - Series B: Biological Sciences. 271, S378-S381.(2004).
Evidence for a South African endemic clade of Ranid frogs. V Congresso Nazionale della Societas Herpetologica Italica.(2004).