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Uyeda, J. C., Drewes R. C., & Zimkus B. M. (2007).  The California Academy of Sciences Gulf of Guinea expeditions (2001,2006) VI. A new species of Phrynobatrachus from the Gulf of Guinea islands and a reanalysis of Phrynobatrachus dispar and P. feae (Anura: Phrynobatrachidae). Proceedings of the California Academy of Sciences. 58, 367-385. Abstract
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